Over the generations their winning behavioural traits would have spread throughout the rest of the population.

For instance, the rate of mutation for behavioural traits has more effects for behavioural mutation within captive populations and some endangered species.

The visual imagery and narratives in his paintings are derived from keen and intelligent observations of social life and behavioural traits.

Behavioural geneticists study the inheritance of behavioural traits.

Multiple genes may simultaneously affect behavioural traits.