knuckle-walkern.

1859 R. Owen Lect. Classif. Mammalia 75 Of the broad-breasted quadrumana, are the knuckle-walkers or the brachiators, i.e. the long-armed gibbons, most nearly and essentially related to the human subject?

1874 J. G. Wood Out of Doors I. 25 When these creatures [the gorilla, chimpansee, and orang-outan] aid their steps by placing the hands on the ground, they have the curious habit of resting the knuckles on the ground..From this peculiarity, the three apes have received the appropriate title of knuckle~walkers.

1967 S. L. Washburn in Proc. Royal Anthropol. Inst. 23/1 Both chimpanzees and gorillas are knuckle walkers.

1973 Nature 10 Aug. 373/2 The first component separates quadrupedal cercopithecoids from both knuckle-walkers (Pan and Gorilla) and the quadrupedal arm-swinger..Ateles.

1967 R. H. Tuttle in Amer. Jrnl. Physical Anthropol. 26 171 Most orang-utans assume one of a variety of flexed hand postures, but they cannot knuckle-walk.

1967 R. H. Tuttle in Amer. Jrnl. Physical Anthropol. 26 171 (title) Knuckle-walking and the evolution of Hominoid hands.

1967 R. H. Tuttle in Amer. Jrnl. Physical Anthropol. 26 171/2 The knuckle-walking posture of chimpanzees and gorillas is unique among primates.

1967 S. L. Washburn in Proc. Royal Anthropol. Inst. 23/1 It appears that our ancestors were arboreal apes for many millions of years, that they then shared a common knuckle-walking stage with the ancestors of the chimpanzee and gorilla, and that only later they became bipeds.

1969 A. H. Schultz Life of Primates v. 56 In the orang-utan the mode of using the hands in walking..is more in the form of a clenched fist, rather awkwardly twisted sideways, and rarely like real knuckle-walking.

1972 O. J. Lewis in R. H. Tuttle Functional & Evolutionary Biol. Primates ix. 212 Thick palmar radiocarpal ligaments are not the exclusive property of knuckle-walkers... It thus seems that knuckle-walking requires no especially striking modifications of the wrist joint.