Onthophagus
Onthophagus
the largest genus of dung beetles of thefamily Scarabaeidae. The males usually have horns or out-growths on the head and often on the pronotum. There areapproximately 1, 500 species. They are found chiefly in the trop-ics; there are about 60 species in the USSR. They dig littleburrows and fill them with dung; the larvae develop in theseburrows. Onthophagus are beneficial as cleaners of the environ-ment and participants in soil formation. Some species are theintermediate hosts of a number of helminths that parasitizedomestic animals.