chemosynthesis

che·mo·syn·the·sis

C0272900 (kē′mō-sĭn′thĭ-sĭs, kĕm′ō-)n. The synthesis of organic compounds by certain bacteria, especially in deep-sea hydrothermal vents, using energy obtained from the chemical oxidation of simple inorganic compounds. Chemosynthesis is thought to have been used by the first forms of life on Earth.

che′mo·syn·thet′ic (-sĭn-thĕt′ĭk) adj.che′mo·syn·thet′i·cal·ly adv.

chemosynthesis

(ˌkɛməʊˈsɪnθɪsɪs) n (Biochemistry) the formation of organic material by certain bacteria using energy derived from simple chemical reactions chemosynthetic adj ˌchemosynˈthetically adv

che•mo•syn•the•sis

(ˌki moʊˈsɪn θə sɪs, ˌkɛm oʊ-)

n. the synthesis of organic compounds within an organism, with chemical reactions providing the energy source. [1900–05] che`mo•syn•thet′ic (-ˈθɛt ɪk) adj. che`mo•syn•thet′i•cal•ly, adv.

che·mo·syn·the·sis

(kē′mō-sĭn′thĭ-sĭs) The formation of organic compounds using the energy released from chemical reactions instead of the energy of sunlight. Bacteria living in deep, dark areas of the ocean are able to survive by chemosynthesis. They use energy derived from the oxidation of inorganic chemicals, such as sulfur from deep volcanic vents, to make their food. Compare photosynthesis.

Thesaurus

Noun

chemosynthesis - synthesis of carbohydrate from carbon dioxide and water; limited to certain bacteria and fungisynthesis - the process of producing a chemical compound (usually by the union of simpler chemical compounds)

chemosynthesis

chemosynthesis,

process in which carbohydrates are manufactured from carbon dioxide and water using chemical nutrients as the energy source, rather than the sunlight used for energy in photosynthesisphotosynthesis
, process in which green plants, algae, and cyanobacteria utilize the energy of sunlight to manufacture carbohydrates from carbon dioxide and water in the presence of chlorophyll. Some of the plants that lack chlorophyll, e.g.
..... Click the link for more information. . Much life on earth is fueled directly or indirectly by sunlight. There are, however, certain groups of bacteria, referred to as chemosynthetic autotrophs, that are fueled not by the sun but by the oxidation of simple inorganic chemicals, such as sulfates or ammonia. Chemosynthetic autotrophs are a necessary part of the nitrogen cyclenitrogen cycle,
the continuous flow of nitrogen through the biosphere by the processes of nitrogen fixation, ammonification (decay), nitrification, and denitrification. Nitrogen is vital to all living matter, both plant and animal; it is an essential constituent of amino acids,
..... Click the link for more information. . Some groups of these bacteria are well suited to conditions that would have existed on the earth billions of years ago, leading some to postulate that these are living representatives of the earliest life on earth. This view has been supported by the discovery of ecosystems that thrive in the hot (350&degC;/660&degF;) water found around hydrothermal vents on the ocean floor. In these ecosystems, the primary producers in the food web are bacteria whose life functions are fueled by inorganic chemicals that seep up from the earth's crust. Other chemosynthetic bacteria are found deep underground, utilizing nitrogen and sulfur compounds as energy sources. See also autotrophautotroph
, in biology, an organism capable of synthesizing its own organic substances from inorganic compounds. Autotrophs produce their own sugars, lipids, and amino acids using carbon dioxide as a source of carbon, and ammonia or nitrates as a source of nitrogen.
..... Click the link for more information. .

Chemosynthesis

a type of nutrition characteristic of certain bacteria whose only carbon source is CO₂ obtained from the energy of oxidation of inorganic compounds.

The discovery of chemosynthesis by S. N. Vinogradskii in 1887 significantly altered prevailing views on the main types of metabolism in living organisms. Unlike photosynthesis, chemosynthesis does not involve the use of light energy but energy derived from oxidation-reductions that must be adequate for the synthesis of adenosine triphosphate (ATP) and that must exceed 10 kcal/mole. Chemosynthetic bacteria are not strictly a taxonomic group; they are classified according to the inorganic substrate they oxidize. They include microorganisms that oxidize hydrogen, carbon monoxide, reduced sulfur compounds, iron, ammonia, nitrites, and antimony.

Hydrogen bacteria are the most numerous and varied group of chemosynthetic organisms. They perform the reaction 6H₂ + 2O₂ + CO₂ = (CH₂O) + 5H₂O, in which (CH₂O) is the conventional designation of the organic substances formed. Compared with other autotrophic microorganisms, hydrogen bacteria have a high growth rate and may produce a large biomass; they are also able to grow on mediums containing organic matter, that is, they are mixotrophic or facultatively chemoautotrophic.

Carboxydation bacteria are similar to hydrogen bacteria, but they oxidize CO by the reaction 25CO + 12O₂ + H₂O + 24CO₂+ (CH₂O). Sulfur bacteria oxidize hydrogen sulfide, thiosulfate, and molecular sulfur to sulfuric acid. Some, for example, Thiobacillus ferrooxidans, oxidize sulfide minerals and ferrous oxide. The capacity of various aquatic sulfur bacteria for chemosynthesis has not yet been demonstrated. Nitrifying bacteria oxidize ammonia to nitrite (first stage of nitrification) and nitrite to nitrate (second stage). Chemosynthesis occurs under anaerobic conditions in some denitrifying bacteria, which oxidize hydrogen or sulfur but often require organic matter for biosynthesis. Chemosynthesis has been described in some strictly anaerobic meth-anogenic bacteria according to the reaction 4H₂ + CO₂ = CH₄ + 2H₂O.

The biosynthesis of organic compounds in chemosynthesis is achieved by autotrophic assimilation of CO₂ (Calvin carbon reduction cycle) just as in photosynthesis. Energy is obtained in the form of ATP from electron transfer via the chain of respiratory enzymes incorporated into the bacterial cell membrane. Some oxidized substances give off electrons to the chain at the cytochrome c level, thereby creating additional energy to synthesize the reducing agent. Owing to the fairly large consumption of energy, chemosynthetic bacteria—with the exception of hydrogen bacteria—produce a small biomass but oxidize a substantial amount of inorganic matter. Since chemosynthetic bacteria control the oxidative parts of the cycle of major elements in the biosphere, they are vitally important in biogeochemistry. Hydrogen bacteria can be used to obtain protein and to free the atmosphere from CO₂ in closed ecological systems. Chemosynthetic bacteria are highly varied morphologically. Although most of them are pseudomonades, they are also found among budding and filamentous bacteria, spirilla, leptospira, and corynebacteria.

REFERENCES

Kuznetsov, S. I. Mikroflora ozer i ee geokhimicheskaia deiatel’nost’. Leningrad, 1970.
Zavarzin, G. A. Litotrofnye mikroorganizmy. Moscow, 1972.
Karavaiko, G. I., S. I. Kuznetsov, and A. I. Golomzik. Rol’ mikroorganizmov v vyshelachivanii metallov iz rud. Moscow, 1972.

G. A. ZAVARZIN

chemosynthesis

[‚kē·mō′sin·thə·səs] (biochemistry) The synthesis of organic compounds from carbon dioxide by microorganisms using energy derived from chemical reactions.

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Words related to chemosynthesis

noun synthesis of carbohydrate from carbon dioxide and water

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